Laboratory Primate Newsletter


2006 Directory of Graduate Programs in Primatology and Primate Research


* Arizona State University, School of Human Evolution and Social Change (formerly Dept. of Anthropology)
PROGRAM DESCRIPTION: PhD in Anthropology (with MA awarded in the process). Within physical anthropology, specializations in primatology are available. Areas of concentration include primate social behavior and ecology, primate positional behavior and functional and evolutionary morphology, primate dental development and life history, and primate evolution. Interdisciplinary training is available in musculoskeletal and neural adaptations in form and function. Facilities include extensive fossil casts and skeletal collections, a variety of specimens for dissection, 3D imaging and analysis capabilities, and excellent computing capabilities. Faculty interests include relationships between social organization and ecology, infant socialization, parental behavior, primate community ecology, and comparative primate functional and evolutionary morphology. Faculty also maintain an association with the Primate Foundation of Arizona, a private chimpanzee breeding colony. Research on chimpanzee social behavior, growth, and development are underway. Financial aid may be available to graduate students on a competitive basis. Aid is in the form of teaching or research assistantships and graduate fellowships.
FACULTY AND THEIR SPECIALTIES: Leanne T. Nash (social behavior and ecology of primates, socialization, nocturnal prosimians, experimental analysis of behavior); Kaye E. Reed (primate community ecology, primate paleoecology, primate evolution, paleoanthropology); Mark A. Spencer (comparative primate functional and evolutionary morphology, biomechanics, cranial evolution, morphometrics, paleoanthropology); Gary T. Schwartz (primate and human evolution, comparative primate dental development, life history).
FOR FURTHER INFORMATION: Drs . Leanne T. Nash [480-965-4812; e-mail:], Mark A. Spencer [480-727-8763; e-mail], Kaye Reed [480-727-6580; e-mail:], or Gary Schwartz [480 967-8684; e-mail], School of Human Evolution and Social Change (formerly Dept of Anthropology), Box 872402, Arizona State University, Tempe, AZ 85287-2402 [480-965-6213; fax: 480-965-7671]; and see <>.

* Primate Foundation of Arizona
PROGRAM DESCRIPTION: A private, non-profit, research institution housing chimpanzees (Pan troglodytes). This group pursues behavioral research with a goal of improving captive management and the well-being of individual animals.
Internships: Behavioral Research Internships and Veterinarian Externships give students in the behavioral and biological sciences the opportunity for research experience. Internships are on a volunteer basis and provide no stipend.
There are three basic components to the Behavioral Research Internship: 1) an introduction to chimpanzee behavior and behavioral observation data collection; 2) chimpanzee psychological wellness program and environmental enrichment training; and 3) research support tasks such as data entry. The introduction to chimpanzee behavioral observation is the primary component of the internship and includes data collection on an assigned project, entering the data into a spreadsheet program, conducting preliminary analyses, and completing a background literature review. Results of the intern project are presented at the end of the internship to the full staff to provide presentation experience.
The Veterinary Student Externship will include working directly with the Attending Veterinarian in the chimpanzee facility.  It will also include scheduled days of working off site in exotic and small animal private veterinary practices.. The basic components of the internship include: 1) an introduction to elements of chimpanzee veterinary care (physical exams, laboratory techniques, and basic pharmacology), and 2) veterinary research training, including the completion of a research project.
Students should have completed at least two years of a four-year program (junior-level standing) in the behavioral or biological sciences. Both undergraduate and graduate students are encouraged to apply. Previous course work and/or experience in primatology/animal behavior is required for all students. Applications are accepted for three internship periods: Summer (June 1 to August 15), Fall (September 1 to November 15), and Spring (March 1 to May 15). Applications should be submitted at least 6 weeks before the desired starting date.
FACULTY AND THEIR SPECIALTIES: Jo Fritz, Director (captive management and behavior); Elaine Videan Ph.D., Research Director (environmental enrichment and well-being, chimpanzee behavior and physiology); Curtis Eng, D.V.M., Attending Veterinarian (Chimpanzee health).
FOR FURTHER INFORMATION: Jo Fritz, Director, Primate Foundation of Arizona, P.O. Box 20027, Mesa, AZ 85277-0027 [480-832-3780; fax 480-830-7039; e-mail].


* University of California, Davis, Anthropology Department
FACULTY AND THEIR SPECIALTIES: Alexander H. Harcourt (primate behavioral ecology); Lynne A. Isbell (primate behavioral ecology and primate evolution); Peter S. Rodman (evolution of primate behavior, behavioral ecology, and primate evolution).
FOR FURTHER INFORMATION: Dept of Anthropology, One Shields Ave, University of California, Davis, CA 95616-8522; or see <>.

* University of California, Davis, Psychology Department
PROGRAM DESCRIPTION: Psychobiology is an area of specialization within the Psychology graduate program.
FACULTY AND THEIR SPECIALTIES: Karen L. Bales (neuroendocrinology of primate social bonding); John P. Capitanio (primate social behavior and development, personality/temperament, psychoneuroimmunology); Richard G. Coss (developmental psychobiology, evolution, experimental aesthetics, antipredator behavior); Leah A. Krubitzer (evolutionary neurobiology); William A. Mason (primate social behavior); Sally P. Mendoza (behavioral endocrinology; physiological basis of primate social relationships, stress, and reproduction); Jeffrey C. Schank (social behavior, individual-based modeling, biorobotics, development).
FOR FURTHER INFORMATION: Graduate Admissions, Department of Psychology, University of California, One Shields Avenue, Davis, CA 95616.


* University of Florida, Psychology Department
FACULTY AND THEIR SPECIALTIES: Marc N. Branch (behavioral pharmacology, experimental analysis of behavior; squirrel monkeys).
FOR FURTHER INFORMATION: Dr. Marc N. Branch, Psychology Dept, University of Florida, Gainesville, FL 32611 [352-392-0601 x205; e-mail:].


* Emory University, Department of Psychology
PROGRAM NAME AND DESCRIPTION: The program in Neuroscience and Animal Behavior (NAB) approaches topics within the areas of neuroscience, physiological psychology, acquired behavior, and ethology as a unified entity.  Thus, the emphasis is on behavior as a biological phenomenon.
FACULTY: Jocelyne Bachevalier, Frans de Waal, David Edwards, Harold Gouzoules, Robert Hampton, Donna Maney, Jack J. McDowell, Darryl Neill, Hillary R. Rodman, Kim Wallen, Michael Zeiler.
FOR FURTHER INFORMATION: Please contact Ms. Terry Legge, Graduate Program Specialist [404-727-7438; e-mail:]; or Dr. Harold Gouzoules, Director of Graduate Studies [404-727-7444; e-mail:]; both at the Department of Psychology, Emory University, Atlanta, GA 30322.

* Georgia Institute of Technology, School of Psychology
PROGRAM NAME AND DESCRIPTION: MS and PhD in Psychology. Program operates in direct conjunction with Zoo Atlanta and the Yerkes National Primate Research Center. A variety of taxonomic groups are studied (carnivores, ungulates, birds, primates), but specialization is in primates.
FACULTY AND THEIR SPECIALTIES: Dr. Terry Maple, Professor (behavior, environmental psychology); Dr. M. Jack Marr, Professor (experimental analysis of behavior); Dr. Mollie Bloomsmith, adjunct professor (behavior, enrichment, well being); Dr. Tara Stoinski, adjunct professor (behavior and cognition).
FOR FURTHER INFORMATION: Dr. Terry Maple, School of Psychology, Georgia Institute of Technology, Atlanta, GA 30332 [e-mail:]; or Dr. Mollie Bloomsmith, Yerkes National Primate Research Center [e-mail:].

* Georgia State University, Language Research Center, Dept of Psychology
PROGRAM DESCRIPTION: The LRC is associated with the graduate program in Social and Cognitive Psychology (with comparative cognition emphasis) and the graduate program in Neuropsychology and Behavioral Neuroscience, as well as with the Brains and Behavior Program involving faculty in biology, psychology, and other departments.
FACULTY AND THEIR SPECIALTIES: Departmental faculty members include David A. Washburn (Director; comparative cognitive psychology), Michael Owren (vocal communication, acoustics and emotion), and E. Sue Savage-Rumbaugh (on leave; language and culture). LRC faculty include Duane M. Rumbaugh (primate intelligence and learning), Charles Menzel (ethology and spatial cognition), Michael Beran (numerical cognition and self-regulation), Emily Klein (animal cognition), and other co-investigators in various disciplines at GSU and other universities.
FOR FURTHER INFORMATION: Language Research Center, Georgia State University, University Plaza, Atlanta, GA 30303-3083 [fax: 404-244-5752; e-mail:]; and see <> or <>.

* University of Georgia, Athens, Psychology Department
PROGRAM NAME: Neuroscience and Behavior with a specialty area in primatology.
FACULTY AND THEIR SPECIALTIES: Irwin S. Bernstein (primatology, social organization, aggression, sex, dominance); Dorothy Fragaszy (primate behavior, cognition, development, motor skills, social behavior). We also enjoy full cooperation with other departments and universities within the University of Georgia system, as well as collaboration with the Yerkes RPRC of Emory University and the Atlanta Zoo.
FOR FURTHER INFORMATION: Neuroscience and Behavior Program, Dept of Psychology, Univ. of Georgia, Athens, GA 30602-3013 [706-542-2174; fax: 706-542-3275]; and see <>.


* Northwestern University Medical School, Department of Cell and Molecular Biology
PROGRAM NAME: Integrated Graduate Program in the Life Sciences.
FACULTY AND THEIR SPECIALTIES: L. R. Cochard (dental allometry); M. Dagosto (prosimian evolution, systematics, locomotion); M. J. Ravosa (experimental functional morphology, skull form); B. T. Shea (growth, allometry, Miocene and recent hominoids).
FOR FURTHER INFORMATION: Any of the above faculty at the Dept of Cell and Molecular Biology, Northwestern Univ. Med. School, 303 E. Chicago Ave, Chicago, IL 60611-3008; and see <>.

* Southern Illinois University, Department of Anthropology
PROGRAM DESCRIPTION: Biological Anthropology: We offer BA, MA, and PhD degrees in Anthropology with a focus on biological anthropology, including primate studies. Primary areas of specialization include general and functional morphology (both dental and skeletal, human and nonhuman primates), and evolution and systematics, particularly of platyrrhines (as well as Eocene/Oligocene primates) and hominoids. We also offer a campus-wide Graduate Certificate in Systematics.
FACULTY AND THEIR SPECIALTIES: Dr. Robert Corruccini (dental anthropology, hominoid and hominid evolution, epidemiology of human populations, statistics); Dr. Susan M. Ford (skeletal anatomy, platyrrhine and early primate evolution and systematics, evolutionary theory, biogeography).
FOR FURTHER INFORMATION: Dept of Anthropology, Southern Illinois Univ., Carbondale, IL 62901-4502 [618-536-6651].

* The University of Chicago, Committee on Evolutionary Biology, Department of Comparative Human Development. 
PROGRAM NAMES: Doctoral programs: Committee on Evolutionary Biology, Department of Comparative Human Development.
FACULTY AND THEIR SPECIALTIES: David Bradley (neuroscience); Nicholas Hatsopoulos (neuroscience); Bruce Lahn (genetics); Wen-Hsiung Li (genetics); Elizabeth Lonsdorf (behavior, ecology); Dario Maestripieri (behavior, ecology); Susan Margulis (behavior, research in zoo settings); Robert D. Martin (evolution); Carole Ober (genetics); Callum Ross (functional morphology, evolution); Russell Tuttle (functional morphology, evolution).
FOR FURTHER INFORMATION: Dario Maestripieri, The University of Chicago, 5730 S. Woodlawn Ave, Chicago, IL 60637 [e-mail:].


* Antioch New England Graduate School, Department of Environmental Studies and Center for Tropical Ecology and Conservation
PROGRAM NAME: Conservation Biology and Environmental Studies.
FACULTY AND THEIR SPECIALTIES: Beth Kaplin, PhD: Specialty: primate seed dispersal behavior, interactions between nonhuman primates and people (use of habitats, crop raiding, hunting), guenon ecology and biogeography, primate conservation.
FOR FURTHER INFORMATION: Antioch New England Graduate School, 40 Avon St., Keene, NH 03431-3516 [603-357-3122; e-mail:].


* City University of New York, Anthropology PhD Program
See under: The New York Consortium in Evolutionary Primatology (NYCEP)

* Columbia University, Ecology, Evolution and Environmental Biology (E3B) Department (also Anthropology Department)
See under: NYCEP

* New York Consortium in Evolutionary Primatology
PROGRAM DESCRIPTION: NYCEP is a graduate training program funded by NSF, most recently by the Integrative Graduate Education and Research Traineeship initiative. It consists of three degree-granting institutions – City University of New York (CUNY), Columbia University (CU), and New York University (NYU) – in collaboration with the American Museum of Natural History (AMNH) and the Wildlife Conservation Society (WCS). Our focus is on nonhuman as well as human primates from the perspectives of comparative morphology, paleontology and systematics, molecular and population genetics, behavior and ecology, and conservation biology. Students in the program take courses in all of these areas at the three universities, attend seminars that draw upon the staff of all five cooperating institutions, and have the opportunity to engage in original research in laboratories, museums, and in the field. Detailed information is available at .
NYCEP is basically an umbrella organization which coordinates course programs and seminars and provides funds for student research and travel support. The graduate programs of the three collaborating universities offer graduate fellowships supported by the IGERT award (up to $30,000 per year for four years, available only to U.S. citizens, nationals and permanent residents), as well as full financial aid programs with regular fellowships and special opportunities for minority students and all highly qualified applicants regardless of nationality. Members of groups underrepresented in science are especially encouraged to apply. Appropriate undergraduate majors for NYCEP applicants include biological anthropology and other life sciences. Students apply to one or more cooperating universities and send a one-page application tracking form (available from the Website) to NYCEP. Annual application deadline is early January.
CORE FACULTY are those with whom students will take most courses and who will be likely dissertation supervisors: Susan Antón, NYU (paleoanthropology, comparative morphology, forensic anthropology); Marina Cords, CU (primate behavior, especially African cercopithecids); Roberto Delgado, CUNY (behavioral ecology, great ape social structure, evolution of human social behavior); Eric Delson, CUNY/AMNH (paleoanthropology; catarrhine systematics and evolution, biochronology); Tony DiFiore, NYU (primate behavior and ecology, population and molecular genetic applications); Todd R. Disotell, NYU (molecular systematics and evolution, catarrhine primates); Terry Harrison, NYU (catarrhine systematics, comparative morphology, primate paleontology); Ralph L. Holloway, CU (paleoneurology, human evolution); Clifford J. Jolly, NYU (genetics, systematics, and comparative morphology of primates); Jeffrey T. Laitman, CUNY (paleoanthropology, evolution of speech); Don J. Melnick, CU (population genetics and molecular evolution of higher primates); John F. Oates, CUNY (ecology and behavior of catarrhine primates, tropical forest conservation); Kate Pechenkina, CUNY (paleopathology, bioarcheology, paleodietary reconstruction); Tom Plummer, CUNY (paleoanthropology, hominid paleontology and paleoecology/behavior, Paleolithic archeology); Alfred Rosenberger, CUNY (evolution of New World monkeys, comparative and functional morphology of dentitions); Vincent Stefan, CUNY (forensic anthropology, human osteology, craniometry); Michael Steiper, CUNY (molecular anthropology, human and other primate genetic adaptations, population genetics, malaria); Sara Stinson, CUNY (population biology of living humans); Larissa Swedell, CUNY (primate, especially cercopithecid, social behavior; population genetics).
RESOURCE FACULTY are available for consultation, may supervise internships and participate on dissertation committees: Walter Bock, CU (vertebrate functional and evolutionary morphology, biomechanics, systematics, evolutionary theory); Tim Bromage, NYU (paleoanthropology and developmental morphology); Peter De Menocal, CU (Pliocene-Pleistocene evolution of tropical climates; African climate and human evolution); Rob De Salle, AMNH (molecular systematics); John Flynn, AMNH (mammalian paleontology and systematics, Primates, Carnivora, South American Cenozoic); Patrick J. Gannon, Mount Sinai/NYU (Primate brain evolution and relationship to communication, neurochemistry); Katerina Harvati, CUNY (also Max-Planck Institute of Evolutionary Anthropology; paleoanthropology, later human evolution and variation, geometric morphometrics); Patrick Hof, Mount Sinai/NYU (neurobiology); Ross D. MacPhee, AMNH (development and systematics of primates and other mammals); Colleen McCann, WCS (conservation biology, behavior and ecology of cercopithecids, hormonal mediation of behavior); Jin Meng, AMNH (paleontology and evolution of early mammals, lagomorphs, and rodents); Juan Carlos Morales, CU (molecular and population genetics, conservation); Michael Novacek, AMNH (systematics of mammals and early primates); Joy Reidenberg, Mount Sinai/NYU (comparative and developmental anatomy of the mammalian skull and upper respiratory tract); John G. Robinson, WCS (conservation biology, Neotropical primates); Robert Rock-well, CUNY (population genetics, population ecology and dynamics, conservation biology); F. James Rohlf, CUNY (also SUNY/Stony Brook; mathematical biology, biostatistics, geometric morphometrics); Mitchell Schaffler, Mount Sinai/NYU (functional and comparative morphology); Eleanor J. Sterling, AMNH (primate social behavior, ecology, and conservation, especially in Madagascar); Katherine St. John, CUNY (computational biology, phylogeny reconstruction and comparison, algorithms); Karyl Swartz, CUNY (comparative psychology, primate cognition); Ian Tattersall, AMNH (systematics and evolution of lemuriform primates and hominids); Herbert S. Terrace, CU (primate language and learning, cognitive psychology); Carl J. Terranova, CUNY (evolutionary anatomy of strepsirrhine primate limbs, developmental and clinical anatomy of human limbs); John A. Van Couvering, CUNY (also Micropaleontology Project; geochronology and stratigraphy of the Old World Cenozoic); John Wahlert, CUNY (mammalian, especially rodent, paleontology, morphology and evolution); Ward Wheeler, AMNH (molecular systematics); Michael Yuan, CU (human dentition, human and other primate endocasts and brain morphology).
FOR FURTHER INFORMATION: Dr. Eric Delson, Dept of Vertebrate Paleontology, American Museum of Natural History, New York, NY 10024 [212-769-5992; fax: 212-769-5842; e-mail:]; or see <>.

* New York University, Anthropology Department
See under: NYCEP


* Duke University, Department of Biological Anthropology and Anatomy
PROGRAM NAME: Graduate Study in Biological Anthropology and Anatomy.
FACULTY AND THEIR SPECIALTIES: Matt Cartmill (anthropoid and primate origins, history of ideas about animal consciousness); Ken Glander (ecology and social organization); William L. Hylander (functional and evolutionary morphology of the masticatory apparatus); Richard F. Kay (anthropoid phylogeny, based especially on cranial and dental anatomy, through paleontological field research); Daniel Schmitt (evolution of locomotion); Elwyn L. Simons (primate paleontology); Kathleen K. Smith (vertebrate evolutionary morphology); John W. Terborgh (tropical forest ecology); Steven Churchill (functional morphology of upper limb bones in later stages of human evolution, Neanderthals); V. Louise Roth (evolutionary modification of growth and development in mammals); Christine Drea (social behavior, social learning, and reproductive endocrinology); Leslie Digby (female strategies and social organization).
FOR FURTHER INFORMATION: Dept. of Biological Anthropology and Anatomy, Director of Graduate Studies, 08 Biological Sciences Bldg, Box 90383, Duke University, Durham, NC 27708.


* Oregon National Primate Research Center
PROGRAM DESCRIPTION: We do not have a formal program in primatology, but we do train pre- and postdoctoral students in using primates for biomedical research. The Oregon National Primate Research Center (ONPRC) is one of eight federally funded centers designed to advance knowledge about human health and disease through research with nonhuman primates. The ONPRC encourages scientists and students to make use of its unique research opportunities in several disciplines, including reproductive biology, neuroscience, perinatal physiology, and immunology and infectious diseases. The Center is an institute of the Oregon Health and Science University (OHSU), and most ONPRC scientists have faculty appointments at the OHSU School of Medicine. The Center staff includes about 60 scientists with PhD, MD, or DVM degrees, as well as 220 technical, support, and service employees. Among the services provided are veterinary care; surgery; pathology; image capture by laser scanning confocal, and electron, microscopy image analysis; molecular and cell biology; radioimmunoassay; flow cytometry; data processing; bibliographic and other library searches; and medical illustration.
FACULTY AND THEIR SPECIALTIES: The scientific expertise of the faculty is focused on molecular and cellular aspects of reproductive biology, neuroscience, and infectious diseases. The Center also employs seven full-time veterinarians who are involved in the daily care of 3200 nonhuman primates and 4000 small laboratory animals.
FOR FURTHER INFORMATION: Oregon National Primate Research Center, 505 N.W. 185th Ave., Beaverton, OR 97006 [503-690-5201]; or visit <>.


* University of Pennsylvania, Departments of Anthropology, Biology, and Psychology
PROGRAM DESCRIPTION: Students may enroll for a PhD with a specialization in primatology in one of the three sponsoring departments; graduate programs will conform in structure and content to the requirements of each department. A group of core interdisciplinary courses is also offered for primatology students, in addition to courses that pertain to their specialties (e.g., cognition, ecology, behavior). Other resources include faculty in ecology and conservation within the Department of Biology; faculty in psycholinguistics and cognitive science in the Department of Psychology and at the Institute for Research in Cognitive Science; and faculty in neuroscience and neuroethology in the Medical School. Cheney and Seyfarth maintain a long-term study of baboons in the Moremi Game Reserve, Okavango Delta, Botswana; in past years their graduate students have also conducted fieldwork in Mexico, Costa Rica, and the Ivory Coast.
FACULTY AND THEIR SPECIALTIES: Dorothy L. Cheney (Biology: behavior, communication, cognition); Robert M. Seyfarth (Psychology: behavior, communication, cognition).
FOR FURTHER INFORMATION: Contact Dr. Cheney or Dr. Seyfarth, Univ. of Pennsylvania, Philadelphia, PA 19104 [e-mail: or].

* University of Pittsburgh, Department of Anthropology
PROGRAM NAME: Physical Anthropology Graduate Program
FACULTY AND THEIR SPECIALTIES: Margaret Judd (human skeletal analysis, paleopathology, bioarcheology); Mark P. Mooney (craniofacial and developmental biology, comparative anatomy, experimental morphology, physiological adaptations to extreme environments, development of animal models for facial clefts); Jeffrey H. Schwartz (method, theory, and philosophy in evolutionary biology; origin and diversification of primates; human and faunal skeletal analysis; dentofacial growth and development); Michael I. Siegel (craniofacial biology, with a clinical specialty in cleft palate; functional anatomy; animal models; physiological adaptation).
FOR FURTHER INFORMATION: Phyllis J. Deasy, Graduate Admissions Coordinator, Department of Anthropology, University of Pittsburgh, Pittsburgh, PA 15260 [e-mail:]; and see <>.

* Bucknell University, Department of Psychology and Program in Animal Behavior
PROGRAM DESCRIPTION: The Bucknell University Animal Behavior Program offers Master’s degrees (MS) in animal behavior. The program does not offer a formal degree in primatology, but primate behavior is an area of specialization offered within the program. Bucknell maintains four social colonies of primates for use in observational studies and noninvasive experiments of behavior and cognition. The Master’s program is designed as an apprenticeship for one or two students to work closely with a sponsoring faculty member.
FACULTY: Dr. Peter G. Judge (specializes in conflict resolution behavior, social cognition, cognitive abilities).
FOR FURTHER INFORMATION: Graduate Studies, Bucknell University, Lewisburg, PA 17837 [570-577-3655]; or see <> or <>.


* University of Texas, Austin, Anthropology Department
PROGRAM DESCRIPTION: MA and PhD degrees are offered in Anthropology, with specialization in physical anthropology, including primate anatomy, ecology, evolution, and behavior.
FACULTY AND THEIR SPECIALTIES: John Kappelman (physical anthropology, paleobiology, primate evolution, functional morphology, stratigraphy; Africa and Asia); Chris Kirk (physical anthropology, primate evolution, functional anatomy and evolution of sensory systems); Liza Shapiro (physical anthropology, primate evolution, functional morphology, locomotion); Deborah Bolnick (physical anthropology, molecular anthropology, ancient DNA).
FOR FURTHER INFORMATION: Dept. of Anthropology, Univ. of Texas, Austin, TX 78712; or see <>.


* Central Washington University, Chimpanzee and Human Communication Institute, Experimental Psychology-Primatology, Dept. of Psychology.
PROGRAM NAME AND DESCRIPTION: MS in Experimental Psychology-Primatology includes opportunity for research in the following areas: chimpanzee language, cognition, and communication.
FACULTY AND THEIR SPECIALTIES: Roger S. Fouts, PhD (chimpanzee language and communication dialects – Psychology); Lori K. Sheeran, PhD (gibbon behavior and conservation – Biological Anthropology); and Mary Lee Jensvold, PhD (chimpanzee sign language studies, conversation repair – Psychology).
FOR FURTHER INFORMATION: Mary Lee Jensvold or Deborah H. Fouts, Chimpanzee and Human Communication Institute, Central Washingon University, Ellensburg, WA 98926-7573 [e-mail: or or]; or see <>.

* University of Washington, Department of Psychology
PROGRAM DESCRIPTION: The Animal Behavior Program at the University of Washington is dedicated to providing the best possible graduate training including research techniques, theory, and investigative work with animals both in the laboratory and in natural habitats, preserves, or progressive zoos. The program leads to the PhD in Psychology, with special training in animal behavior (including primate social behavior). It is administered by the core faculty in animal behavior, who are listed below. One of the great assets of this Animal Behavior Program is the interest and competence of faculty in departments other than Psychology. Cordial and cooperative relationships exist with behavior-oriented colleagues in Zoology, Biology, Anthropology, Wildlife Science (College of Fisheries and School of Forest Resources), the Conservation Biology Program, the Neurobiology Program, the Regional Primate Research Center, and the Human Development and Disabilities Center’s interdepartmental pathway in primatology. Excellent rapport and research affiliations also exist with the Woodland Park Zoological Gardens, Point Defiance Zoo, the Seattle Aquarium, Northwest Trek, the Friday Harbor biology and marine research laboratories, and colleagues in the greater Puget Sound area.
FACULTY AND THEIR SPECIALTIES: Michael D. Beecher, (animal communication, avian sociobiology and ecology); Gene P. Sackett (primate development and behavior); David P. Barash (sociobiology, behavioral ecology, animal behavior and evolution); Eliot A. Brenowitz (avian behavior, neuroethology, neuroendocrinology, animal communication); Sean O’Donnell (social behavior, especially of insects; evolution of eusociality, particularly division of labor and task allocation; behavioral genetics; and physiology); Ellen Covey (comparative neural bases, anatomy, physiology, function, and modeling of audition; auditory-motor pathways; echolocation; and auditory temporal patterns and processing networks). Also available to facilitate student projects are James Ha (DNA studies in animal behavior), Randall Kyes (Indonesian macaque field site), and Julie Worlein (primate AIDS research), all of whom are graduate faculty with primary appointments at the Regional Primate Center.
FOR FURTHER INFORMATION: Michael D. Beecher, PhD, Dept. of Psychology, Box 351525, University of Washington, Seattle, WA 98l95-1525 [e-mail:].


* Wisconsin National Primate Research Center, University of Wisconsin-Madison, Graduate School. supported by a base operating grant from the National Institutes of Health-National Center for Research Resources
PROGRAM DESCRIPTION: The research program at the WNPRC has opportunities for graduate studies in several areas, especially reproductive and developmental biology, including placental biology and stem cell research, immunogenetics, virology and AIDS vaccine development, veterinary medicine, aging and metabolic disease, psychology and psychobiology, and neurobiology, including Parkinson’s disease research.
Students may conduct research at the WNPRC by enrolling in an appropriate academic department at the University of Wisconsin-Madison and choosing a faculty advisor with WNPRC affiliation. Current faculty have appointments in various departments in the Medical School, College of Letters and Science, School of Veterinary Medicine, and College of Agriculture and Life Sciences, as well as such interdisciplinary programs as the Endocrinology-Reproductive Physiology Program, Interdepartmental Graduate Program in Nutritional Sciences, the Biology of Aging and Age-Related Diseases Training Program, and the Neuroscience Training Program. For information about these departments and programs, potential students should write to The Graduate School, Bascom Hall, UW-Madison, Madison, WI 53706; or visit <>.
FACULTY AND STAFF: The WNPRC supports the work of approximately 75 doctoral-level scientists on campus and approximately 150 affiliates based at other academic institutions. There are 24 faculty and staff on the WNPRC Executive Committee.
FOR FURTHER INFORMATION: Joseph W. Kemnitz, Director, WNPRC, 1220 Capitol Ct, Madison, WI 53715-1299. Director’s Office and general information: [608-263-3500; fax: 608-265-2067]; or see: <>.

· University of Wisconsin, Milwaukee, Department of Anthropology
PROGRAM DESCRIPTION: Behavioral endocrinology, parasitology, behavioral and evolutionary ecology, population genetics, demography, DNA analysis for paternity determination, evolution, behavior, comparative anatomy and functional morphology of humans and nonhuman primates. The Department of Anthropology has graduate programs leading to MS and PhD degrees.
FACULTY AND THEIR SPECIALTIES: Fred Anapol (primate functional morphology, muscle biology, skeletal analysis [Adjunct, Department of Biological Sciences]); Michael P. Muehlenbein (primate reproductive ecology, behavioral endocrinology, ecological parasitology); Trudy R. Turner (DNA analysis, nonhuman primate population genetics, ecology and evolution, medical genetics); Neil C. Tappen, emeritus (primate anatomy, ecology, and evolution; structure and function of bone and muscle). In the Department of Biological Sciences: R. J. Hutz (regulation of ovarian function in monkeys, effects of xenobiotics on estrogen receptor signaling); Andrew J. Petto (anatomy and physiology; animal models for research and education; demography and evolutionary ecology).
FOR FURTHER INFORMATION: Department of Anthropology, University of Wisconsin-Milwaukee, Milwaukee, WI 53201 [414-229-4175]; or see <>.


* Australian National University, Canberra, School of Archaeology and Anthropology
PROGRAM DESCRIPTION: MA (by course work and thesis), MPhil (by thesis alone) and PhD programs in Biological Anthropology, including primatology.
The PhD consists solely of research; no course work is involved. Graduates of this program have worked on such topics as primate digestive strategies; European Miocene hominoids; gibbon social organization and ecology in central Borneo; human cranial thickness; behavioural adaptations of snare-injured chimpanzees in Uganda; palaeogeography and mammalian biogeography in Southeast Asia; and Central American spider monkey ecology and taxonomy.  The Physical Anthropology Laboratory of the School of Archaeology and Anthropology has a collection of primate skulls and skeletons, Australian mammal skulls, and casts of fossil primates including hominins. Students from overseas wishing to study at Australian Universities are charged a Foreign Students’ Fee, currently $A19,200; there are a few Overseas Student Scholarships which cover this fee. Further scholarships are available to cover living expenses.
FACULTY AND THEIR SPECIALTIES: Colin P. Groves (primate taxonomy, evolution, functional morphology, behavior, ecology); Robert Attenborough (behavior, genetics, epidemiology); Marc Oxenham (skeletal biology, palaeopathology).
FOR FURTHER INFORMATION: Professor C. P. Groves, School of Archaeology & Anthropology, Australian National University, Canberra, ACT 0200, Australia.


* Pontifícia Universidade Católica do Rio Grande Do Sul, Faculdade de Biociências
PROGRAM NAME: MSc & PhD Program in Zoology
FACULTY AND THEIR SPECIALTIES: Julio Cesar Bicca-Marques (ecology, behavior, and conservation of New World monkeys).
FOR FURTHER INFORMATION: Secretaria do Programa de Pós-Graduação em Biociências, Av. Ipiranga 6681 – Prédio 12C Sala 254, 90619-900 Porto Alegre, RS, Brasil [Phone/Fax (51) 3320-3568; e-mail: or].


* University of Calgary, Department of Anthropology
PROGRAM DESCRIPTION: Master’s and Doctoral programs available in primate studies, principally field-based and with a behavioral ecology approach. Both MA and PhD programs require course work, a formal research proposal defense, field research minimum of 4 and 12 months respectively, preparation and defense of a thesis, and candidacy and second-language exams at the doctoral level.  The department has research relationships with the Monkey River, Belize, site at which an annual field school is conducted; the Boabeng Fiema Monkey Sanctuary in Ghana, at which an annual field school is conducted; Santa Rosa National Park in Costa Rica; and various field sites in Madagascar.
FACULTY AND THEIR SPECIALTIES: Linda Fedigan (life histories, behavioral ecology and conservation of polygamous primates; field sites in Costa Rica and Japan); Steig Johnson (behavioral ecology, biogeography, speciation, and conservation biology of lemurs, especially brown lemurs; field sites in Madagascar); Mary Pavelka (aging and reproduction; social relationships; behavioral ecology; Japanese macaques; Belizean black howlers); Pascale Sicotte (social relationships; colobine socioecology; ape socioecology; male reproductive competition).
FOR FURTHER INFORMATION: Dept. of Anthropology, Univ. of Calgary, Calgary, Alberta T2N 1N4, Canada [e-mail: or or or]; or see: <>.


* Universities of St Andrews, Stirling, Edinburgh, and Abertay
PROGRAM NAME AND DESCRIPTION: Scottish Primate Research Group (SPRG). The SPRG was formed in 1987, with a core membership of fieldworkers from the triangle of Edinburgh, St Andrews, and Stirling Universities, each about an hour’s travelling time from the others. Regular joint research meetings and seminars by national and international visitors are held and a network of associates swells attendance at these meetings. Field studies by core Group members are carried out at several sites in Africa, Asia, and South America; studies of captive primates rely on well-housed breeding groups at Edinburgh Zoo, as well as major primate centres in France, Japan and the U.S.A. The focus of SPRG research is the natural behaviour, mentality and ecology of primates. Results are often of a kind that informs welfare and conservation policies, and members of the SPRG do not conduct invasive research.
FACULTY AND THEIR SPECIALTIES: James R. Anderson (Psychology, Stirling: social behavior, learning and cognition, environmental enrichment); Hannah M. Buchanan-Smith (Psychology, Stirling: color vision, welfare, polyspecific associations); Richard Byrne (Psychology, St Andrews: cognition in primates and other animals); Christine Caldwell (Psychology, Stirling: learning and cognition, the evolution of culture); Tecumseh Fitch (Psychology, St Andrews: primate vocal communication and the evolution of language); Juan-Carlos Gomez (Psychology, St Andrews: communication, joint attention, theory of mind); Scott Hardie (Psychology, Abertay: social behavior of New World primates); Kevin N. Laland (Biology, St Andrews: social learning, cultural transmission and innovation); Phyllis Lee (Psychology, Stirling: social and physical development, social evolution, reproduction, conservation); Sarah Vick (Psychology, Stirling: social cognition and the evolution of communication, facial expressions, eye gaze); Alexander Weiss (Psychology, Edinburgh: the evolution and behavior genetics of personality and subjective well-being in primates, especially great apes and humans); Andrew Whiten (Psychology, St Andrews: social learning, culture and cognition); Klaus Zuberbuhler (Psychology, St Andrews: communication in African primates).
FOR FURTHER INFORMATION: Those interested in applying for a postgraduate position should write to “Postgraduate Admissions” at the relevant institution: School of Psychology, University of St Andrews, St Andrews, Fife KY16 9JP, Scotland <>; Department of Psychology, University of Stirling, Stirling FK9 4LA, Scotland <>; Department of Psychology, 7 George Square, Edinburgh, EH8 9JZ, Scotland <>; or Dr. S. Hardie, Division of Psychology, Abertay University, Marketgait House, Marketgait, Dundee DD1 1NG, Scotland <>; and see: <>.


* National Sun Yat-sen University, Department of Biological Sciences.
PROGRAM DESCRIPTION: Students are encouraged to study the ecology, social behavior, demographic patterns, and reproduction of Formosan macaques, Macaca cyclopis, leading to Master’s and Doctoral degrees. Long-term field studies on Formosan macaques have been carried out at the Mt. Longevity study site at Kaohsiung for over a decade.
FACULTY AND THEIR SPECIALTIES: Minna J. Hsu (life history parameters, sexual selection, reproductive strategies and behavioral ecology of macaques; field sites in Taiwan and India); and Govindasamy Agoramoorthy (behavioral ecology, sociobiology and population studies of monkeys and apes; field sites in Borneo, Taiwan, and India).
FOR FURTHER INFORMATION: Dept of Biological Sciences, National Sun Yat-sen University, Kaohsiung 80424, Taiwan [e-mail: or].

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Cover photograph of a cotton-top tamarin (Saguinus oedipus) at Roger Williams Park Zoo, by Mark Abbott

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Last updated: December 19, 2005