Laboratory Primate Newsletter


Directory of Graduate Programs in Primatology and Primate Research (2004)


* Arizona State University, Anthropology Department
PROGRAM DESCRIPTION: PhD in Anthropology (with MA awarded in the process). Within physical anthropology, specializations in primatology are available. Areas of concentration include primate social behavior and ecology, primate positional behavior and functional and evolutionary morphology, and primate evolution. Interdisciplinary training is available in musculoskeletal and neural adaptations in form and function. Facilities include extensive fossil casts and skeletal collections, a variety of specimens for dissection, 3D imaging and analysis capabilities, and excellent computing capabilities. Faculty interests include relationships between social organization and ecology, infant socialization, parental behavior, primate community ecology, and comparative primate functional and evolutionary morphology. Faculty also maintain an association with the Primate Foundation of Arizona, a private chimpanzee breeding colony. Research on chimpanzee social behavior, growth, and development are underway. Financial aid may be available to graduate students on a competitive basis. Aid is in the form of teaching or research assistantships and graduate fellowships.
FACULTY AND THEIR SPECIALTIES: Leanne T. Nash (social behavior and ecology of primates, socialization, nocturnal prosimians, experimental analysis of behavior); Mary W. Marzke (comparative primate functional morphology and evolutionary morphology, human evolution, growth and development); Kaye E. Reed (primate community ecology, primate paleoecology, primate evolution, paleoanthropology).
FOR FURTHER INFORMATION: Drs. Leanne T. Nash, Mary W. Marzke, or Kaye Reed, Dept of Anthropology, Box 872402, Arizona State University, Tempe, AZ 85287-2402 [480-965-6213; fax: 480-965-7671; Dr. Nash: 480-965-4812; e-mail:; Dr. Marzke: 480-965-6237; e-mail:; Dr. Reed: 480-727-6580; e-mail:]; and see <>.

* Primate Foundation of Arizona
PROGRAM DESCRIPTION: A private, non-profit, chimpanzee (Pan troglodytes) colony pursuing behavioral research with a goal of improving captive management and the well-being of individual animals. Internships: Behavioral Research Internships provide college students in the behavioral and biological sciences the opportunity for behavioral research experience. There are three basic components: 1) an introduction to chimpanzee behavior and behavioral observation data collection; 2) chimpanzee psychological wellness program and environmental enrichment training; and 3) research support tasks such as data entry. Introduction to chimpanzee behavioral observation is the primary component of the internship and includes data collection on an assigned project, entering the data into a spreadsheet program, conducting preliminary analyses, and completing a background literature review. Results of the intern project are presented at the end of the internship to the full staff to provide presentation experience. Internships are on a volunteer basis and provide no stipend. Students should have completed at least two years of a four-year program (junior-level standing) in the behavioral or biological sciences. Both undergraduate and graduate students are encouraged to apply. Previous course work and/or experience in primatology/animal behavior is required for all students. Applications are accepted for three internship periods: Summer (June 1 to August 31), Fall (September 1 to November 28), and Spring (March 1 to May 30). Applications should be submitted at least 6 weeks before the desired starting date.
FACULTY AND THEIR SPECIALTIES Jo Fritz, Director (captive management and behavior); Elaine Videan M.S., Research Director (environmental enrichment and well-being, chimpanzee behavior).
FOR FURTHER INFORMATION Jo Fritz, Director, Primate Foundation of Arizona, P.O. Box 20027, Mesa, AZ 85277-0027 [480-832-3780; fax 480-830-7039; e-mail].


* California State University, San Marcos, Department of Psychology
PROGRAM NAME: Master of Arts in General Experimental Psychology.
FACULTY AND THEIR SPECIALTIES: Nancy Caine (callitrichid behavior), with possibilities for collaboration with primatologists at the San Diego Zoo and San Diego Wild Animal Park.
FOR FURTHER INFORMATION: Nancy Caine, Dept. of Psychology, CSU San Marcos, San Marcos, CA 92096 [e-mail:].

* University of California, Davis, Anthropology Department
FACULTY AND THEIR SPECIALTIES: Alexander H. Harcourt (primate behavioral ecology); Lynne A. Isbell (primate behavioral ecology); Peter S. Rodman (evolution of primate behavior, behavioral ecology, and primate evolution).
FOR FURTHER INFORMATION: Dept of Anthropology, One Shields Ave, University of California, Davis, CA 95616-8522; or see <>.

* University of California, Davis, Psychology Department
PROGRAM DESCRIPTION: Psychobiology is an area of specialization within the Psychology graduate program.
FACULTY AND THEIR SPECIALTIES: John P. Capitanio (primate social behavior and development, personality/temperament, psychoneuroimmunology); Richard G. Coss (developmental psychobiology, evolution, experimental aesthetics, antipredator behavior); Leah A. Krubitzer (evolutionary neurobiology); William A. Mason (primate social behavior); Sally P. Mendoza (behavioral endocrinology, physiological basis of primate social relationships, stress, and reproduction); Jeffrey C. Schank (social behavior, individual-based modeling, development).
FOR FURTHER INFORMATION: Graduate Admissions, Department of Psychology, University of California, One Shields Avenue, Davis, CA 95616.


* University of Florida, Psychology Department
FACULTY AND THEIR SPECIALTIES: Marc N. Branch (behavioral pharmacology, experimental anal-ysis of behavior; squirrel monkeys).
FOR FURTHER INFORMATION: Dr. Marc N. Branch, Psychology Dept, University of Florida, Gainesville, FL 32611 [352-392-0601 x205; e-mail:].


* Emory University, Department of Psychology
PROGRAM NAME AND DESCRIPTION: The program in Neuroscience and Animal Behavior (NAB) approaches topics within the areas of neuroscience, physiological psychology, acquired behavior, and ethology as a unified entity.
FACULTY: Frans de Waal, David Edwards, Harold Gouzoules, Donna Maney, Jack J. McDowell, Darryl Neill, Hillary R. Rodman, Kim Wallen, Michael Zeiler.
FOR FURTHER INFORMATION: Please contact Ms. Terry Legge, Graduate Program Specialist [404-727-7438; e-mail:]; or Dr. Harold Gouzoules, Director of Graduate Studies [404-727-7444; e-mail:]; both at the Department of Psychology, Emory University, Atlanta, GA 30322.

* Georgia Institute of Technology, Psychology Department
PROGRAM NAME AND DESCRIPTION: MS and PhD in Psychology. Program operates in direct conjunction with Zoo Atlanta. A variety of taxonomic groups are studied (carnivores, ungulates, birds, primates), but specialization is in primates.
FACULTY AND THEIR SPECIALTIES: Dr. Terry Maple, Professor (behavior, environmental psychology); Dr. M. Jack Marr, Professor (experimental analysis of behavior); Dr. Mollie Bloomsmith, adjunct professor (behavior, enrichment, well being); Dr. Tara Stoinski, adjunct professor (behavior and cognition).
FOR FURTHER INFORMATION: Dr. Terry Maple, School of Psychology, Georgia Institute of Technology, Atlanta, GA 30332 [e-mail:]; or Dr. Mollie Bloomsmith, Zoo Atlanta, 800 Cherokee Ave, Atlanta, GA 30315 [e-mail:].

* Georgia State University, Language Research Center, Dept of Psychology or Dept of Biology
PROGRAM DESCRIPTION: Social/Cognitive (with comparative cognition emphasis) in Psychology; bio-behavioral, cognitive, and language studies with pri-mates in Biology.
FACULTY AND THEIR SPECIALTIES: Departmental faculty include David A. Washburn (Director; comparative cognitive psychology) and E. Sue Savage-Rumbaugh (PI for culture and communication; biopsychology, primatology, apes and language). LRC faculty include Duane M. Rumbaugh (primate intelligence and cognition), Charles Menzel (ethology and spatial cognition), Claudio Cantalupo (comparative neuropsychology), Michael Beran (numerical cognition), and other co-investigators in various disciplines at GSU and other universities.
FOR FURTHER INFORMATION: Language Research Center, Georgia State University, University Plaza, Atlanta, GA 30303-3083 [fax: 404-244-5752; e-mail:]; <>; or <>.

* University of Georgia, Athens, Psychology Department
PROGRAM NAME: Neuroscience and Behavior with a specialty area in primatology.
FACULTY AND THEIR SPECIALTIES: Irwin S. Bernstein (primatology, social organization, aggression, sex, dominance); Dorothy Fragaszy (primate behavior, cognition, development, motor skills, social behavior). We also enjoy full cooperation with other departments and universities within the University of Georgia system, as well as collaboration with the Yerkes RPRC of Emory University and the Atlanta Zoo.
FOR FURTHER INFORMATION: Neuroscience & Behavior Program, Dept of Psychology, Univ. of Georgia, Athens, GA 30602-3013 [706-542-2174; fax: 706-542-3275]; and see <>


* Northwestern University Medical School, Department of Cell and Molecular Biology
PROGRAM NAME: Integrated Graduate Program in the Life Sciences.
FACULTY AND THEIR SPECIALTIES: L. R. Cochard (dental allometry); M. Dagosto (prosimian evolution, systematics, locomotion); M. J. Ravosa (experimental functional morphology, skull form); B. T. Shea (growth, allometry, Miocene and recent hominoids).
FOR FURTHER INFORMATION: Any of the above faculty at the Dept of Cell and Molecular Biology, Northwestern Univ. Med. School, 303 E. Chicago Ave, Chicago, IL 60611-3008; and see <>.

* Southern Illinois University, Department of Anthropology
PROGRAM DESCRIPTION: Biological Anthropology: We offer BA, MA, and PhD degrees in Anthropology with a focus on biological anthropology, including primate studies. Primary areas of specialization include general and functional morphology (both dental and skeletal, human and nonhuman primates), and evolution and systematics, particularly of platyrrhines (as well as Eocene/Oligocene primates) and hominoids. We also offer a campus-wide Graduate Certificate in Systematics.
FACULTY AND THEIR SPECIALTIES: Dr. Robert Corruccini (dental anthropology, hominoid and hominid evolution, epidemiology of human populations, statistics); Dr. Susan M. Ford (skeletal anatomy, platyrrhine and early primate evolution and systematics, evolutionary theory, biogeography); Dr. Christopher Stojanowski (human osteology, bioarchaeology, paleogenetics, southeastern U.S.).
FOR FURTHER INFORMATION: Dept of Anthropology, Southern Illinois Univ., Carbondale, IL 62901-4502 [618-536-6651].

* The University of Chicago, Dept. of Anthropology, Dept. of Ecology and Evolution, Dept. of Organismal Biology and Anatomy, Dept. of Psychology, Committee on Evolutionary Biology, Committee on Human Development
PROGRAM NAMES: Doctoral programs: Committee on Evolutionary Biology, Committee on Human Development, Department of Anthropology, Department of Ecology and Evolution, Department of Organismal Biology and Anatomy, Department of Psychology.
FACULTY AND THEIR SPECIALTIES: David Bradley (Psychology: vision and neuroscience); Dario Maestripieri (Human Development; Evolutionary Biology: behavior, development, evolution); Sue Margulis (Evolutionary Biology: behavior, reproduction, research in zoo settings); Robert D. Martin (Evolutionary Biology: evolution, behavior, reproduction, genetics, ecology and conservation); Martha McClintock (Psychology; Evolutionary Biology; Human Development: menstrual synchrony, hormones, pheromonal communication); Callum Ross (Organismal Biology and Anatomy: primate anatomy and functional morphology; behavior and evolution); Russell Tuttle (Anthropology; Evolutionary Biology: primate morphology, locomotion, and behavior); Leigh Van Valen (Evolutionary Biology: population biology and evolutionary theory); Carole Ober (Human Genetics: genetics).
FOR FURTHER INFORMATION: Dario Maestripieri, The University of Chicago, 5730 S. Woodlawn Ave, Chicago, IL 60637 [e-mail:].


* Antioch New England Graduate School, Department of Environmental Studies and Center for Tropical Ecology and Conservation
PROGRAM NAME: Conservation Biology and Environmental Studies.
FACULTY AND THEIR SPECIALTIES: Beth Kaplin, PhD: Specialty: primate seed dispersal behavior, interactions between nonhuman primates and people (use of habitats, crop raiding, hunting), guenon ecology and biogeography, primate conservation.
FOR FURTHER INFORMATION: Antioch New England Graduate School, 40 Avon St., Keene, NH 03431-3516 [603-357-3122; e-mail:].


* City University of New York, Anthropology PhD Program
See under: The New York Consortium in Evolutionary Primatology

* Columbia University, Ecology, Evolution and Environmental Biology (E3B) Department (also Anthropology Department)
See under: The New York Consortium in Evolutionary Primatology

* New York Consortium in Evolutionary Primatology (NYCEP)
PROGRAM DESCRIPTION: NYCEP is a graduate training program funded by NSF, most recently by the Integrative Graduate Education and Research Traineeship initiative. It consists of three degree-granting institutions - City University of New York (CUNY), Columbia University (CU), and New York University (NYU) - in collaboration with the American Museum of Natural History (AMNH) and the Wildlife Conservation Society (WCS). Our focus is on nonhuman as well as human primates from the perspectives of comparative morphology, paleontology and systematics, molecular and population genetics, behavior and ecology, and conservation biology. Students in the program take courses in all of these areas at the three universities, attend seminars that draw upon the staff of all five cooperating institutions, and have the opportunity to engage in original research in laboratories, museums, and in the field. Detailed information is available at <>.
NYCEP is basically an umbrella organization which coordinates course programs and seminars and provides funds for student research and travel support. The graduate programs of the three collaborating universities offer graduate fellowships supported by the IGERT award (up to $27,500 for four years), as well as full financial aid programs with regular fellowships and special opportunities for minority students and all highly qualified applicants regardless of nationality. Members of groups underrepresented in science are especially encouraged to apply. Appropriate undergraduate majors for NYCEP applicants include biological anthropology and other life sciences. Students apply to one or more cooperating universities and send a one-page application tracking form (available from the Website) to NYCEP. Annual application deadline is early January.
CORE FACULTY are those with whom students will take most courses and who will be likely dissertation supervisors: Susan Antón, NYU (paleoanthropology, comparative morphology, forensic anthropology); Tim Bromage, CUNY (paleoanthropology and developmental morphology); Marina Cords, CU (primate behavior, especially African cercopithecids); Eric Delson, CUNY (paleoanthropology; catarrhine systematics and evolution, biochronology); Tony DiFiore, NYU (primate behavior and ecology, population and molecular genetic applications); Todd R. Disotell, NYU (molecular systematics and evolution, catarrhine primates); Terry Harrison, NYU (catarrhine systematics, comparative morphology, and primate paleontology); Katerina Harvati, NYU (paleoanthropology, later human evolution and variation, geometric morphometrics); Ralph L. Holloway, CU (paleoneurology, human evolution); Clifford J. Jolly, NYU (genetics, systematics, and comparative morphology of primates); Jeffrey T. Laitman, CUNY (paleoanthropology, evolution of speech); Don J. Melnick, CU (population genetics and molecular evolution of higher primates); John F. Oates, CUNY (ecology and behavior of catarrhine primates, tropical forest conservation); Tom Plummer, CUNY (paleoanthropology, hominid paleontology and paleoecology/behavior, Paleolithic archeology); Vincent Stefan, CUNY (forensic anthropology, human osteology, craniometry); Sara Stinson, CUNY (population biology of living humans); Karyl Swartz, CUNY (comparative psychology, primate cognition); Larissa Swedell, CUNY (primate, especially cercopithecid, social behavior; population genetics).
RESOURCE FACULTY are available for consultation, may supervise internships and participate on dissertation committees: Walter Bock, CU (vertebrate functional and evolutionary morphology, biomechanics, systematics, evolutionary theory); Roberto Delgado, CUNY (behavioral ecology, great ape social structure, evolution of human social behavior); Rob De Salle, AMNH (molecular systematics); Patrick J. Gannon, Mount Sinai/NYU (Primate brain evolution and relationship to communication, neurochemistry); Patrick Hof, Mount Sinai/NYU (neurobiology); Cathi Lehn, AMNH (primate genetics, conservation); Ross D. MacPhee, AMNH (development and systematics of primates and other mammals); Colleen McCann, WCS (conservation biology, behavior and ecology of cercopithecids, hormonal mediation of behavior); Jin Meng, AMNH (paleontology and evolution of early mammals, lagomorphs, and rodents); Juan Carlos Morales, CU (molecular and population genetics, conservation); Ken Mowbray, AMNH (comparative, developmental and functional morphology, human craniology, paleoanthropology); Michael Novacek, AMNH (systematics of mammals and early primates); Kate Pechenkina, CUNY (paleopathology, bioarcheology, paleodietary reconstruction); David Reddy, AMNH (computer visualization, morphometrics); Joy Reidenberg, Mount Sinai/NYU (comparative and developmental anatomy of the mammalian skull and upper respiratory tract); John G. Robinson, WCS (conservation biology, Neotropical primates); Robert Rockwell, CUNY (population genetics, population ecology and dynamics, conservation biology); F. James Rohlf, CUNY (also SUNY/Stony Brook) (mathematical biology, biostatistics, geometric morphometrics); Alfred Rosenberger, CUNY (evolution of New World monkeys, comparative and functional morphology of dentitions); Mitchell Schaffler, Mount Sinai/NYU (functional and comparative morphology); Michael Steiper, CUNY (molecular anthropology, human and other primate genetic adaptations, population genetics, malaria); Eleanor J. Sterling, AMNH (primate social behavior, ecology, and conservation, especially in Madagascar); Katherine St. John, CUNY (computational biology, phylogeny reconstruction and comparison, algorithms); Ian Tattersall, AMNH (systematics and evolution of lemuriform primates and hominids); Carl J. Terranova, CUNY (evolutionary anatomy of strepsirrhine primate limbs, developmental and clinical anatomy of human limbs); John A. Van Couvering, AMNH (geochronology and stratigraphy of the Old World Cenozoic); John Wahlert, CUNY (mammalian, especially rodent, paleontology, morphology and evolution); Ward Wheeler, AMNH (molecular systematics); Michael Yuan, CU (human dentition, human and other primate endocasts and brain morphology).
FOR FURTHER INFORMATION: Dr. Eric Delson, Dept of Vertebrate Paleontology, American Museum of Natural History, New York, NY 10024 [212-769-5992; fax: 212-769-5842; e-mail:]; or see <>.

* New York University, Anthropology Department
See under: The New York Consortium in Evolutionary Primatology


* Duke University, Department of Biological Anthropology and Anatomy
PROGRAM NAME: Graduate Study in Biological Anthropology and Anatomy.
FACULTY AND THEIR SPECIALTIES: Matt Cartmill (anthropoid and primate origins, history of ideas about animal consciousness); Kenneth E. Glander (ecology and social organization); William L. Hylander (functional and evolutionary morphology of the masticatory apparatus); Richard F. Kay (anthropoid phylogeny, based especially on cranial and dental anatomy, through paleontological field research); Theresa R. Pope (interrelationship between social organization, behavioral ecology, and genetic structure of primate populations); Elwyn L. Simons (primate paleontology); Kathleen K. Smith (vertebrate evolutionary morphology); John W. Terborgh (tropical forest ecology); Steven Churchill (functional morphology of upper limb bones in later stages of human evolution, Neanderthals); V. Louise Roth (evolutionary modification of growth and development in mammals); Christine Drea (social behavior, social learning, and reproductive endocrinology); Diane Brockman (reproductive ecology and endocrinology); Leslie Digby (female strategies and social organization).
FOR FURTHER INFORMATION: Dept. of Biological Anthropology and Anatomy, Director of Graduate Studies, 08 Biological Sciences Bldg, Box 90383, Duke University, Durham, NC 27708.


* Miami University, Department of Zoology
PROGRAM DESCRIPTION: Master’s and PhD degrees in Zoology, specializing in primatology. Strong links to Biological Anthropology (which has no graduate program). No nonhuman primates on campus, but connections to local zoos. (Ohio is the only state to have two breeding colonies of Pan paniscus, at Cincinnati and Columbus Zoos.) Focus on ethology and ecology of anthropoids in Africa.
FACULTY AND THEIR SPECIALTIES: Linda F. Marchant (affiliate in Anthropology: laterality of hand function, chimpanzee behavior, videography); William C. McGrew (laterality of hand function, cultural primatology, ape behavioral ecology).
FOR FURTHER INFORMATION: Graduate Admissions, Dept. of Zoology, Miami Univ., Oxford, OH 45056 [513-529-3100; fax: 513-529-6900]; and see <>.


* Oregon National Primate Research Center
PROGRAM DESCRIPTION: We do not have a formal program in primatology, but we do train pre-and postdoctoral students in using primates for biomedical research. The Oregon National Primate Research Center (ONPRC) is one of eight federally funded centers designed to advance knowledge about human health and disease through research with nonhuman primates. The ONPRC encourages scientists and students to make use of its unique research opportunities in several disciplines, including reproductive biology, neuroscience, perinatal physiology, and immunology and infectious diseases. The Center is an institute of the Oregon Health and Science University (OHSU), and most ONPRC scientists have faculty appointments at the OHSU School of Medicine. The Center staff includes about 60 scientists with PhD, MD, or DVM degrees, as well as 220 technical, support, and service employees. Among the services provided are veterinary care, surgery, pathology, image capture by laser scanning confocal, and electron, microscopy image analysis, molecular and cell biology, radioimmunoassays, flow cytometry, data processing, bibliographic and other library searches, and medical illustration.
FACULTY AND THEIR SPECIALTIES: The scientific expertise of the faculty is focused on molecular and cellular aspects of reproductive biology, neuroscience, and infectious diseases. The Center also employs seven full-time veterinarians who are involved in the daily care of 3200 nonhuman primates and 4000 small laboratory animals.
FOR FURTHER INFORMATION: Oregon National Primate Research Center, 505 N.W. 185th Ave., Beaverton, OR 97006 [503-690-5301].


* University of Pennsylvania, Departments of Anthropology, Biology, and Psychology
PROGRAM DESCRIPTION: Students may enroll for a PhD with a specialization in primatology in one of the three sponsoring departments; graduate programs will conform in structure and content to the requirements of each department. A group of core interdisciplinary courses is also offered for primatology students, in addition to courses that pertain to their specialty (e.g., cognition, ecology, behavior). Other resources include faculty in ecology and conservation within the Department of Biology; faculty in psycholinguistics and cognitive science in the Department of Psychology and at the Institute for Research in Cognitive Science; and faculty in neuroscience and neuroethology in the Medical School. Cheney and Seyfarth maintain a long-term study of baboons in the Moremi Game Reserve, Okavango Delta, Botswana; in past years their graduate students have also conducted fieldwork in Mexico, Costa Rica, and the Ivory Coast.
FACULTY AND THEIR SPECIALTIES: Dorothy L. Cheney (Biology: behavior, communication, cognition); Robert M. Seyfarth (Psychology: behavior, communication, cognition).
FOR FURTHER INFORMATION: Contact Dr. Cheney or Dr. Seyfarth, Univ. of Pennsylvania, Philadelphia, PA 19104 [e-mail: or].

* University of Pittsburgh, Department of Anthropology
PROGRAM NAME: Physical Anthropology Graduate Program
FACULTY AND THEIR SPECIALTIES: Mark P. Mooney (craniofacial and developmental biology, comparative anatomy, experimental morphology, physiological adaptations to extreme environments, development of animal models for facial clefts); Jeffrey H. Schwartz (method, theory, and philosophy in evolutionary biology; origin and diversification of primates; human and faunal skeletal analysis; dentofacial growth and development); Michael I. Siegel (craniofacial biology, with a clinical speciality in cleft palate; functional anatomy; animal models; physiological adaptation).
FOR FURTHER INFORMATION: Phyllis J. Deasy, Graduate Admissions Coordinator, Department of Anthropology, University of Pittsburgh, Pittsburgh, PA 15260 [e-mail:]; and see <>.

* Bucknell University, Department of Psychology and Program in Animal Behavior
PROGRAM DESCRIPTION: The Bucknell University Animal Behavior Program offers Master's degrees (MS) in animal behavior. The program does not offer a formal degree in primatology, but primate behavior is an area of specialization offered within the program. Bucknell maintains four social colonies of primates for use in observational studies and noninvasive experiments of behavior and cognition. The Master's program is designed as an apprenticeship for one or two students to work closely with a sponsoring faculty member.
FACULTY: Dr. Peter G. Judge (specializes in conflict resolution behavior, social cognition, cognitive abilities).
FOR FURTHER INFORMATION: Graduate Studies, Bucknell University, Lewisburg, PA 17837 [570-577-3655]; or see <>. TEXAS

* University of Texas, Austin, Anthropology Department
PROGRAM DESCRIPTION: MA and PhD degrees are offered in Anthropology, with specialization in physical anthropology, including primate anatomy, ecology, evolution, and behavior.
FACULTY AND THEIR SPECIALTIES: John Kappelman (physical anthropology, paleobiology, primate evolution, functional morphology, stratigraphy; Africa and Asia); E. Christopher Kirk (physical anthropology, primate sensory systems and cranio-dental morphology); Liza Shapiro (physical anthropology, primate evolution, functional morphology, locomotion).
FOR FURTHER INFORMATION: Dept. of Anthropology, University of Texas, Austin, TX 78712; or see <>.


* Central Washington University, Chimpanzee and Human Communication Institute, Experimental Psychology-Primatology, Dept. of Psychology.
PROGRAM NAME AND DESCRIPTION: MS in Experimental Psychology-Primatology includes opportunity for research in the following areas: chimpanzee language, cognition, communication, and post-conflict interaction.
FACULTY AND THEIR SPECIALTIES: Roger S. Fouts, PhD (chimpanzee language and communication dialects - Psychology), Lori K. Sheeran, PhD (gibbon behavior and conservation - Biological Anthropology), and Mary Lee Jensvold, PhD (chimpanzee sign language studies - conversation repair - Psychology).
FOR FURTHER INFORMATION: Dr. Roger S. Fouts, Chimpanzee and Human Communication Institute, Central Washingon University, Ellensburg, WA 98926-7573 [e-mail: or]; or see <>.

* University of Washington, Department of Psychology
PROGRAM DESCRIPTION: The Animal Behavior Program at the University of Washington is dedicated to providing the best possible graduate training including research techniques, theory, and investigative work with animals both in the laboratory and in natural habitats, preserves, or progressive zoos. The program leads to the PhD in Psychology, with special training in animal behavior (including primate social behavior). It is administered by the core faculty in animal behavior, who are listed below. One of the great assets of this Animal Behavior Program is the interest and competence of faculty in departments other than Psychology. Cordial and cooperative relationships exist with behavior-oriented colleagues in Zoology, Biology, Anthropology, Wildlife Science (College of Fisheries and School of Forest Resources), the Conservation Biology Program, the Neurobiology Program, the Regional Primate Research Center, and the Human Development and Disabilities Center’s interdepartmental pathway in primatology. Excellent rapport and research affiliations also exist with the Woodland Park Zoological Gardens, Point Defiance Zoo, the Seattle Aquarium, Northwest Trek, the Friday Harbor biology and marine research laboratories, and colleagues in the greater Puget Sound area.
FACULTY AND THEIR SPECIALTIES: Joan S. Lockard (primate social behavior, human ethology, sociobiology, zoo animal behavior, neurobehavior); Michael D. Beecher, (animal communication, avian sociobiology and ecology); Gene P. Sackett (primate development and behavior); David P. Barash (sociobiology, behavioral ecology, animal behavior and evolution); Eliot A. Brenowitz (avian behavior, neuroethology, neuroendocrinology, animal communication); Sean O’Donnell (social behavior, especially of insects; evolution of eusociality, particularly division of labor and task allocation; behavioral genetics; and physiology); Ellen Covey (comparative neural bases, anatomy, physiology, function, and modeling of audition; auditory-motor pathways; echolocation; and auditory temporal patterns and processing networks). Also available to facilitate student projects are James Ha (DNA studies in animal behavior), Randall Kyes (Indonesian macaque field site), and Julie Worlein (primate AIDS research), all of whom are graduate faculty with primary appointments at the Regional Primate Center.
FOR FURTHER INFORMATION: Joan S. Lockard, PhD, Dept. of Psychology, Box 351525, University of Washington, Seattle, WA 98l95-1525 [e-mail:].


* National Primate Research Center, University of Wisconsin-Madison, Graduate School
PROGRAM DESCRIPTION: The research program at the WPRC has opportunities for graduate studies in several areas, especially reproductive and developmental biology, immunogenics and vaccine development, aging, neurobiology, and biogerontology. Students may conduct research at the WPRC by enrolling in an appropriate academic department at the University of Wisconsin-Madison and choosing a faculty advisor with WPRC affiliation. Current faculty have appointments in various departments in the Medical School, College of Letters and Science, School of Veterinary Medicine, and College of Agriculture and Life Sciences, as well as such interdisciplinary programs as the Endocrinology-Reproductive Physiology Program, Interdepartmental Graduate Program in Nutritional Sciences, the Biology of Aging and Age-Related Diseases Training Program, and the Neuroscience Training Program. For information about these departments and programs, potential students should write to The Graduate School, Bascom Hall, UW-Madison, Madison, WI 53706; or visit <>.
FACULTY AND THEIR SPECIALTIES: The WPRC has approximately 50 doctoral-level scientists on campus and approximately 150 affiliates based at other academic institutions. Faculty on the WPRC Executive Committee and their academic departments are: Joseph W. Kemnitz, Director, Physiology; David H. Abbott, Obstetrics and Gynecology; Christopher Coe, Psychology; Thaddeus Golos, Obstetrics and Gynecology; Ei Terasawa, Pediatrics; James Thomson, Anatomy; David Watkins, Pathology; Richard Weindruch, Medicine (Geriatrics).
FOR FURTHER INFORMATION: Joseph W. Kemnitz, Director, WPRC, 1220 Capitol Ct, Madison, WI 53715-1299. Director's Office and general information: [608-263-3500; fax: 608-265-2067]; or see: <>.

* University of Wisconsin, Milwaukee, Department of Anthropology
PROGRAM DESCRIPTION: Ecology, population genetics, comparative anatomy, and aging in primates, especially African monkeys. DNA analysis for paternity determination of nonhuman primates. Evolution, behavior, and functional morphology of nonhuman primates. The Department of Anthropology has graduate programs leading to MS and PhD degrees.
FACULTY AND THEIR SPECIALTIES: Fred Anapol (primate functional morphology, muscle biology, skeletal analysis [Adjunct, Department of Biological Sciences]); Trudy R. Turner (DNA analysis, nonhuman primate population genetics, ecology and evolution, medical genetics); Neil C. Tappen, emeritus (primate anatomy, ecology, and evolution; structure and function of bone and muscle). In the Department of Biological Sciences: R. J. Hutz (regulation of ovarian function in monkeys, effects of xenobiotics on estrogen receptor signaling).
FOR FURTHER INFORMATION: Dept. of Anthropology, University of Wisconsin-Milwaukee, Milwaukee, WI 53201.


* Australian National University, Canberra, School of Archaeology and Anthropology
PROGRAM DESCRIPTION: MA (by course work and thesis), MPhil (by thesis alone) and PhD programs in Biological Anthropology, including primatology. The PhD consists solely of research; no course work is involved. Graduates of this program have worked on colobine dentition, primate digestive strategies, Southeast Asian macaque variation, European Miocene hominoids, and gibbon social organization and ecology in central Borneo. The Physical Anthropology Laboratory of the Department of Archaeology and Anthropology has a collection of primate skulls and skeletons, Australian mammal skulls, and casts of fossil primates including hominids. Students from overseas wishing to study at Australian Universities are charged a Foreign Students’ Fee, currently A$13,500 (or, for a lab-based PhD, A$17,000); there are a few Overseas Student Scholarships which cover this fee. Further scholarships are available to cover living expenses.
FACULTY AND THEIR SPECIALTIES: Colin P. Groves (primate taxonomy, evolution, functional morphology, behavior, ecology); Robert Attenborough (behavior, genetics, epidemiology); Marc Oxenham (skeletal biology, palaeopathology). Collaboration is also possible with Simon Easteal (John Curtin School of Medical Research, same university), specializing in primate genetics, including DNA.
FOR FURTHER INFORMATION: Professor C. P. Groves, School of Archaeology & Anthropology, Australian National University, Canberra, ACT 0200, Australia.


* Pontifícia Universidade Católica do Rio Grande Do Sul, Faculdade de Biociências
PROGRAM NAME: MSc & PhD Program in Zoology
FACULTY AND THEIR SPECIALTIES: Julio Cesar Bicca-Marques (ecology, behavior, and conservation of New World monkeys).
FOR FURTHER INFORMATION: Secretaria do Programa de Pós-Graduação em Biociências, Av. Ipiranga 6681 - Prédio 12C Sala 254, 90619-900 Porto Alegre, RS, Brasil [Phone/Fax (51) 3320-3568; e-mail:].


* University of Calgary, Department of Anthropology
PROGRAM DESCRIPTION: Master's and Doctoral programs available in primatological studies, principally oriented towards behavioral and behavioral ecology approaches. Both programs require course work, a formal research proposal defense, a candidacy examination for doctoral students, field research minimum of 4 and 12 months respectively, and preparation and defense of a thesis. The department has research relationships with various primate research centers and zoos in the USA; the Monkey River, Belize site at which an annual field school is conducted; Boabeng Fiema Monkey Sanctuary in Ghana; and other field sites.
FACULTY AND THEIR SPECIALTIES: Linda Fedigan (life histories, sexual selection, and behavioral ecology of monkeys living in multi-male, multi-female societies; field sites in Costa Rica, Japan, and the U.S.; research on gender and science); Mary McDonald Pavelka (aging and reproduction; social relationships; behavioral ecology; Japanese macaques; Belizean black howlers); Pascale Sicotte (social relationships, colobine socioecology, ape socioecology, male reproductive competition).
FOR FURTHER INFORMATION: Dept of Anthropology, Univ. of Calgary, 2500 University Dr. NW, Calgary, Alberta T2N 1N4 Canada [e-mail: or or]; or see <>.


* University of Liverpool Hominid Palaeontology Research Group (Department of Human Anatomy and Cell Biology) and School of Archaeology, Classics and Oriental Studies (Department of Archaeology)
PROGRAM DESCRIPTION: MSc in Early Hominid Studies. An intensive, interdisciplinary course over one year provides a broadly based theoretical and practical understanding of our own origins and biology and that of our closest relatives within the larger context of climatic change and the evolution of life. It provides an excellent basis for further research in the field. Graduates with a first degree in a variety of arts and sciences subjects may enroll.
FACULTY AND THEIR SPECIALTIES: Robin Crompton (primate ecology, behavior, and evolution); Robin Dunbar (primate social behavior and evolution); Michael Günther (functional morphology and biomechanics); John Gowlett (paleolithic archeology; early hominid sites; radiocarbon dating); Alf Latham (geochronology and geoarcheology); Gabriele Macho (early hominid evolution; gnathic and dental evolution, function, and development); John Shaw (paleomagnetism); Anthony Sinclair (archeological theory; late paleolithic).
FOR FURTHER INFORMATION: Gabriele Macho, Hominid Palaeontology Research Group, Dept of Human Anatomy and Cell Biology, Univ. of Liverpool, P.O. Box 147, Liverpool L69 3BX, England [e-mail:].

* University of Surrey Roehampton, U.K.
PROGRAM DESCRIPTION: One-year Master of Research (MRes) degree program. This program provides a unique opportunity to study primate biology in depth. It teaches original research and places findings into a theoretical context, providing preparation for advanced research (PhD and consultancy work). It combines theoretical investigation with laboratory and field work on a range of topics. Practical investigations will be carried out in zoos, local habitats, museums and laboratories. After the first semester the emphasis will be on independent research, with all students carrying out an in-depth piece of original research. This will be written up as a dissertation and a paper in a form suitable for publication in a peer-reviewed scientific journal. Key areas of study will be: · Ecology and behavior: Methods used in surveying and gathering biological information; methods of recording behavior in the field. · Diet and foraging: Observing and investigating behavioral and physical dietary adaptations; field and laboratory techniques for gathering data; analyzing nutritional and foraging data from wild and captive primates. · Life-history evolution: Allometry; reproductive life history variables; comparative analysis of life-history and brain size evolution. · Reproduction: Laboratory techniques for gathering data and analyzing reproductive hormone data in wild and captive primates; the evolution of mating strategies. · Zoos and museums as a resource for the study of primates; the ethics of studying captive primates. · Methods of analyzing physical and behavioral adaptations (e.g. locomotion, sensory systems); phylogenetic reconstructions and interpretations of adaptations.
FOR FURTHER INFORMATION: School of Life and Sport Sciences, University of Surrey Roehampton, West Hill, London SW15 3SN, England [020 8392 3524; e-mail:]; or see <>.


* Universities of St Andrews, Stirling, Edinburgh, and Abertay
PROGRAM NAME AND DESCRIPTION: Scottish Primate Research Group (SPRG). The SPRG. was formed in 1987, with a core membership of fieldworkers from the triangle of Edinburgh, St Andrews and Stirling Universities, each about an hour's travel from the others. Regular joint research meetings and seminars by national and international visitors are held and a network of associates swells attendance at these meetings. Field studies by core Group members are carried out at several sites in Africa, Asia and South America; studies of captive primates rely on well-housed breeding groups at Edinburgh and Belfast Zoos, as well as major primate centers in France and U.S.A. The focus of SPRG research is the natural behavior, mentality, and ecology of primates. Results are often of a kind that inform welfare and conservation policies, and members of the SPRG do not conduct invasive research.
FACULTY AND THEIR SPECIALTIES: James R. Anderson (Psychology, Stirling: social behavior, learning and cognition; environmental enrichment); Hannah Buchanan-Smith (Psychology, Stirling: color vision, welfare); Richard Byrne (Psychology, St Andrews: cognition in primates, manual skill and laterality, foraging behavior); Tecumseh Fitch (Psychology, St Andrews: primate vocal communication and the evolution of language); Juan-Carlos Gomez (Psychology, St Andrews: communication, joint attention, theory of mind); Scott Hardie (Psychology, Abertay: social behavior of New World primates); Kevin N. Laland (Biology, St Andrews; social learning, cultural transmission and innovation); Natalie Waran (Veterinary Clinical Studies, Edinburgh: welfare of animals in captive conditions); Andrew Whiten (Psychology, St Andrews: social learning, culture and cognition); Klaus Zuberbuhler (Psychology, St Andrews: communication in African primates).
FOR FURTHER INFORMATION: Postgraduate Admissions, School of Psychology, Univ. of St Andrews, St Andrews, Fife, KY16 9JU, Scotland; or Dept of Psychology, Univ. of Stirling, Stirling, FK9 4LA, Scotland, or Dr. S. Hardie, Division of Psychology, Abertay University, Marketgait House, Marketgait, Dundee DD1 1NG, Scotland; and see <>.


* National Sun Yat-sen University, Department of Biological Sciences, Kaohsiung 80424, Taiwan.
PROGRAM DESCRIPTION: Students are encouraged to study the ecology, social behavior, demographic patterns, and reproduction of Formosan macaques, Macaca cyclopis, leading to Master’s and Doctoral degrees. Long-term field studies on Formosan macaques have been carried out at the Mt. Longevity study site at Kaohsiung for over a decade.
FACULTY AND THEIR SPECIALTIES: Minna J. Hsu (life history parameters, sexual selection, reproductive strategies and behavioral ecology of macaques; field sites in Taiwan and India); and Govindasamy Agoramoorthy (behavioral ecology, sociobiology and population studies of monkeys and apes; field sites in Borneo, Taiwan, and India).
FOR FURTHER INFORMATION: Dept of Biological Sciences, National Sun Yat-sen University, Kaohsiung 80424, Taiwan [e-mail: or].

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All correspondence concerning the Newsletter should be addressed to:
Judith E. Schrier, Psychology Department, Box 1853, Brown University
Providence, Rhode Island 02912. [401-863-2511; FAX: 401-863-1300]
e-mail address:

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The Newsletter is supported by U. S. Public Health Service Grant RR-00419 from the Comparative Medicine Program, National Center for Research Resources, N.I.H.

Cover illustration is a Chinese print, purchased in a “tourist shop”. The inscription reads something like “Monkey, Garden, Pleasure”

Copyright (c) 2004 by Brown University

Last updated: December 15, 2003