Laboratory Primate Newsletter


2010 Directory of Graduate Programs in Primatology and Primate Research


* Arizona State University, School of Human Evolution and Social Change
PROGRAM DESCRIPTION: PhD in Anthropology (with MA awarded in the process). Within physical anthropology, specializations in primatology are available. Areas of concentration include primate social behavior and ecology, primate positional behavior and functional and evolutionary morphology, primate dental development and life history, and primate evolution. Interdisciplinary training is available in musculoskeletal and neural adaptations in form and function. Facilities include extensive fossil casts and skeletal collections, a variety of specimens for dissection, 3D-imaging and analysis capabilities, and excellent computing capabilities. Faculty interests include relationships between social organization and ecology, infant socialization, parental behavior, primate community ecology, and comparative primate functional and evolutionary morphology. Financial aid may be available to graduate students on a competitive basis. Aid is in the form of teaching or research assistantships and graduate fellowships.
FACULTY AND THEIR SPECIALTIES: Leanne T. Nash (social behavior and ecology of primates, socialization, nocturnal prosimians, experimental analysis of behavior); Kaye E. Reed (primate community ecology, primate paleoecology, primate evolution, paleoanthropology); Mark A. Spencer (comparative primate functional and evolutionary morphology, biomechanics, cranial evolution, morphometrics, paleoanthropology); and Gary T. Schwartz (primate and human evolution, comparative primate dental development, life history).
FOR FURTHER INFORMATION: Drs. Leanne T. Nash [480-965-4812; e-mail: [email protected]], Mark A. Spencer [480-727-8763; e-mail [email protected]], Kaye Reed [480-727-6580; e-mail: [email protected]], or Gary Schwartz [480 967-8684; e-mail [email protected]], School of Human Evolution and Social Change (formerly Dept of Anthropology), Box 872402, Arizona State University, Tempe, AZ 85287-2402 [480-965-6213; fax: 480-965-7671]; and see <>.


* University of California, Davis, Anthropology Department
FACULTY AND THEIR SPECIALTIES: Alexander H. Harcourt (primate behavioral ecology, biogeography; conservation); Lynne A. Isbell (primate behavioral ecology, primate origins and evolution); Andrew J. Marshall (community ecology, tropical forest ecology; primate conservation); and David G. Smith (primate genetics).
FOR FURTHER INFORMATION: Dept of Anthropology, One Shields Ave, University of California, Davis, CA 95616-8522; or see <>.

* University of California, Davis, Psychology Department
PROGRAM DESCRIPTION: Psychobiology is an area of specialization within the Psychology graduate program.
FACULTY AND THEIR SPECIALTIES: Karen L. Bales (neuroendocrinology of primate social bonding); John P. Capitanio (primate social behavior and development, personality/temperament, psychoneuroimmunology); Richard G. Coss (developmental psychobiology, evolution, experimental aesthetics, antipredator behavior); Leah A. Krubitzer (evolutionary neurobiology); William A. Mason (primate social behavior); Sally P. Mendoza (behavioral endocrinology; physiological basis of primate social relationships, stress, and reproduction); Jeffrey C. Schank (social behavior, individual-based modeling, biorobotics, development); and Brian C. Trainor (behavioral endocrinology; genes and behavior).
FOR FURTHER INFORMATION: Graduate Admissions, Dept of Psychology, University of California, One Shields Ave, Davis, CA 95616.


* University of Georgia, Athens, Psychology Department
PROGRAM NAME: Neuroscience and Behavior with a specialty area in primatology.
FACULTY AND THEIR SPECIALTIES: Irwin S. Bernstein (primatology, social organization, aggression, sex, dominance) and Dorothy Fragaszy (primate behavior, cognition, development, motor skills, social behavior). We also enjoy full cooperation with other departments and universities within the University of Georgia system, as well as collaboration with the Yerkes RPRC of Emory University and the Atlanta Zoo.
FOR FURTHER INFORMATION: Neuroscience and Behavior Program, Dept of Psychology, Univ. of Georgia, Athens, GA 30602-3013 [706-542-2174; fax: 706-542-3275]; and see <>.


* American Museum of Natural History, Richard Gilder Graduate School
See under: The New York Consortium in Evolutionary Primatology (NYCEP)

* City University of New York, Anthropology PhD Program
See under: NYCEP

* Columbia University, Department of Ecology, Evolution & Environmental Biology
See under: NYCEP

* New York Consortium in Evolutionary Primatology
PROGRAM DESCRIPTION: NYCEP is a graduate training program funded by the National Science Foundation’s Integrative Graduate Education and Research Traineeship (IGERT) initiative. It consists of four degree-granting institutions – City University of New York (CUNY), the American Museum of Natural History (AMNH), Columbia University (CU), and New York University (NYU) – in collaboration with the Wildlife Conservation Society (WCS). Our focus is on nonhuman as well as human primates from the perspectives of comparative morphology, paleontology and systematics, molecular and population genetics, behavior and ecology, and conservation biology. Students in the program take courses in all of these areas at the three universities and AMNH, attend seminars that draw upon the staff of all five cooperating institutions, and have the opportunity to engage in original research in laboratories, museums, and in the field. Detailed information is available at <>. NYCEP provides funds for research and travel support and coordinates course programs and seminars.
NYCEP is basically an umbrella organization which coordinates course programs and and provides funds for student research and travel support. The graduate programs of the four collaborating institutions offer graduate fellowships supported by the IGERT award (up to $30,000 per year for up to four years), as well as full financial aid programs with regular fellowships and special opportunities for minority students. IGERT funds can only be awarded to U.S. citizens and nationals, but highly qualified applicants regardless of nationality may be supported in some institutions. Members of groups underrepresented in science are especially encouraged to apply. Appropriate undergraduate majors for NYCEP applicants include biological anthropology and other life (or earth) sciences. Students apply to one or more cooperating universities and send a one-page tracking form to NYCEP; this is available from the Website or from Dr. Delson (see below). Annual application deadline is early January.
FACULTY AND THEIR SPECIALTIES: CORE FACULTY are those with whom students will take most courses and who will be likely dissertation supervisors: Susan Antón, NYU (paleoanthropology, comparative morphology, forensic anthropology); Shara Bailey, NYU (paleoanthropology, dental anthropology); Doug Boyer, CUNY (primate evolutionary and functional morphology, paleontology, paleoanthropology, systematics and evolution); Marina Cords, CU (primate behavior, especially African cercopithecids); Eric Delson, CUNY (paleoanthropology, catarrhine systematics and evolution, biochronology, morphometrics); Tony DiFiore, NYU (primate behavior and ecology, population and molecular genetic applications); Todd R. Disotell, NYU (molecular systematics and evolution, catarrhine primates); John Flynn, AMNH (Neotropical paleontology, including primates; carnivoran evolution); Chris Gilbert, CUNY (paleoanthropology, primate [especially cercopithecid] comparative morphology, paleontology and evolution); William Harcourt-Smith, CUNY (paleoanthropology, evolution of bipedalism, primate locomotion, morphometrics); Terry Harrison, NYU (catarrhine systematics, comparative morphology, and primate paleontology); Ralph L. Holloway, CU (paleoneurology, human evolution); Jeffrey T. Laitman, CUNY (paleoanthropology, evolution of speech); Don J. Melnick, CU (population genetics and molecular evolution of higher primates); Kate Pechenkina, CUNY (paleopathology, bioarcheology, paleodietary reconstruction); Tom Plummer, CUNY (paleoanthropology, hominid paleontology and paleoecology/behavior, Paleolithic archeology); Herman Pontzer, CUNY (locomotor biomechanics and energetics, foraging ecology, hominid evolution); Ryan Raaum, CUNY (population genetics, human molecular variation, phylogeography); Alfred Rosenberger, CUNY (evolution of New World monkeys, comparative and functional morphology of dentitions); Jessica Rothman, CUNY (African primate ecology and behavior, nutrition, evolutionary ecology); Vincent Stefan, CUNY (forensic anthropology, human osteology, craniometry); Michael Steiper, CUNY (molecular anthropology, human and other primate genetic adaptations, population genetics, malaria); Sara Stinson, CUNY (population biology of living humans); Larissa Swedell, CUNY (primate, especially cercopithecid, social behavior; population genetics).
RESOURCE FACULTY, some from other institutions, are available for consultation, and may supervise internships and participate on dissertation committees: Gail Ashley, Rutgers (Quaternary geology, stratigraphy, geochronology); Marie-Pierre Aubry, Rutgers (Cenozoic chronostratigraphy, micropaleontology); Bill Berggren, Rutgers (Cenozoic chronostratigraphy, micropaleontology); Walter Bock, CU (vertebrate functional and evolutionary morphology, biomechanics, systematics, evolutionary theory); Tim Bromage, NYU Dental School (paleoanthropology and developmental morphology); Nick Conard, Tübingen (European Middle and Upper Paleolithic archeology and art); Peter deMenocal, CU (geochemistry; African climate and human evolution); Rob De Salle, AMNH (molecular systematics); Craig Feibel, Rutgers (African Plio-Pleistocene geochronology, stratigraphy, climate); Patrick J. Gannon, Hofstra and Mount Sinai/NYU (primate brain evolution and relationship to communication; neurochemistry); Katerina Harvati, Tübingen and CUNY (paleoanthropology, later human evolution and variation, geometric morphometrics); Sidney Hemming, CU (geochronology); Mike Hickerson, CUNY (ecological modeling, phylogeography); Patrick Hof, Mount Sinai/NYU (neurobiology); Jean-Jacques Hublin, Max-Planck Inst, Leipzig (paleoanthropology); Clifford J. Jolly, NYU (genetics, systematics, and comparative morphology of primates); Ross D. MacPhee, AMNH (development and systematics of primates and other mammals); Colleen McCann, WCS (conservation biology, behavior and ecology of cercopithecids, hormonal mediation of behavior); Shannon McPherron, Max-Planck Inst, Leipzig (European and North African Middle and Upper Paleolithic archeology, site formation); Jin Meng, AMNH (mammalian paleontology and evolution, China); Jason Munshi-South, CUNY (ecology and conservation genetics of primates and other mammals of SE Asia); Xijun Ni, AMNH and CUNY (paleontology and evolution of early primates and other mammals, China); Michael Novacek, AMNH (systematics of mammals and early primates); Joy Reidenberg, Mount Sinai/NYU (comparative and developmental anatomy of the mammalian skull and upper respiratory tract); John G. Robinson, WCS (conservation biology, Neotropical primates); Robert Rockwell, CUNY (population genetics, population ecology and dynamics, conservation biology); F. James Rohlf, CUNY and SUNY/Stony Brook (mathematical biology, biostatistics, geometric morphometrics); Eric Sanderson, WCS (GIS approaches in conservation science); Mitchell Schaffler, CCNY/CUNY (functional and comparative morphology); Eleanor J. Sterling, AMNH (primate social behavior, ecology, and conservation, especially in Madagascar); Katherine St. John, CUNY (computational biology, phylogeny reconstruction and comparison, algorithms); Carl Swisher, Rutgers (geochronology and stratigraphy of primate-bearing deposits); Ian Tattersall, AMNH (systematics and evolution of lemuriform primates and hominids); Herb Terrace, CU (comparative psychology); Christian Tryon, NYU (African Lower and Middle Paleolithic archeology, geochronology, geoarcheology); John A. Van Couvering, Micropaleontology Project (geochronology and stratigraphy of the Old World Cenozoic); Linda Vigilant, Max-Planck Inst, Leipzig (primate molecular and conservation genetics); John Wahlert, CUNY (mammalian, especially rodent, paleontology, morphology and evolution); Ward Wheeler, AMNH (molecular systematics); Randall White, NYU (European Middle and Upper Paleolithic archeology, ornamentation).
FOR FURTHER INFORMATION: Dr. Eric Delson, Dept of Vertebrate Paleontology, American Museum of Natural History, New York, NY 10024 [212-769-5992; fax: 212-769-5842; e-mail: [email protected]]; or see <>.

* New York University, Anthropology Department See under: NYCEP


* Duke University, Evolutionary Anthropology (formally Biological Anthropology and Anatomy)
PROGRAM DESCRIPTION: Undergraduate and PhD programs in Evolutionary Anthropology including: Primate Behavior and Ecology, Primate Paleontology; Cognitive Evolution; and Functional Morphology.
FACULTY AND THEIR SPECIALTIES: Steven Churchill (functional morphology, hominin paleontology); Leslie Digby (primate behavior, reproductive competition, lemurs); Christine Drea (social behavior, social learning, reproductive endocrinology); Ken Glander (ecology and social organization); Brian Hare (cognitive evolution, human and non-human apes); Richard F. Kay (primate phylogeny, paleontological field research); Anne Pusey (behavioral ecology, parent-offspring interaction, sex differences in development, dispersal patterns, mating systems); Daniel Schmitt (evolution of primate locomotor mechanics); Christine Wall (evolutionary and functional morphology of primate skull); and Blythe Williams (primate paleontology; functional morphology).
FOR FURTHER INFORMATION: Dept. of Evolutionary Anthropology, Director of Graduate Studies, 08 Biological Sciences Bldg, Box 90383, Duke University, Durham, NC 27708; [DGS Assistant: Lisa Squires: 919-684-4124; e-mail: [email protected]].


* Bucknell University, Program in Animal Behavior
PROGRAM DESCRIPTION: The Bucknell University Animal Behavior Program offers Master’s degrees (MS) in animal behavior. The program does not offer a formal degree in primatology, but primate behavior is an area of specialization offered within the program. Bucknell maintains four colonies of socially living primates for use in observational studies and noninvasive experiments of behavior and cognition. The Master’s program is designed as an apprenticeship for one or two students to work closely with a sponsoring faculty member.
FACULTY: Dr. Peter G. Judge (specializes in conflict resolution behavior, social cognition, cognitive abilities).
FOR FURTHER INFORMATION: Graduate Studies, Bucknell University, Lewisburg, PA 17837; 570-577-3655. Also see: <> or <>.

* University of Pennsylvania, Departments of Anthropology, Biology, and Psychology
PROGRAM DESCRIPTION: Students may enroll for a PhD with a specialization in primatology in one of the three sponsoring departments; graduate programs will conform in structure and content to the requirements of each department. A group of core interdisciplinary courses is also offered for primatology students, in addition to courses that pertain to their specialties (e.g., cognition, ecology, behavior). Other resources include faculty in ecology and conservation within the Department of Biology; faculty in psycholinguistics, cognitive science, animal behavior, and the evolution of human behavior in the Department of Psychology and at the Institute for Research in Cognitive Science; faculty in human biology, primatology, and endocrinology in the Department of Anthropology; and faculty in neuroscience and neuroethology in the Medical School. Dorothy Cheney and Robert Seyfarth accept only graduate students who plan to do field research, and in past years have supervised students working on elephants in Sri Lanka, hyenas in Kenya, rhesus monkeys on Cayo Santiago, and gelada baboons in Ethiopia. Eduardo Fernandez-Duque is particularly interested in students who will consider working on neotropical monogamous species in Ecuador or Argentina. Claudia Valeggia's research interests include primate reproductive ecology (primarily human), reproductive endocrinology, maternal and child health, and health of indigenous populations in Latin America. Tad Schurr is interested in molecular anthropology, modern human evolution, primate biological variation, biomedical genetics, and ancient DNA research.
FACULTY AND THEIR SPECIALTIES: Dorothy L. Cheney (Biology: behavior, communication, cognition); Robert M. Seyfarth (Psychology: behavior, communication, cognition); Eduardo Fernandez-Duque (Anthropology: behavior, ecology, conservation); Claudia Valeggia (Anthropology: energetics, endocrinology, reproduction); and Tad Schurr (Anthropology: genetics).
FOR FURTHER INFORMATION: Contact Dr. Cheney [e-mail: [email protected]] or Dr. Seyfarth [e-mail: [email protected]] or Dr. Fernandez-Duque [e-mail: [email protected]], University of Pennsylvania, Philadelphia, PA 19104.


* University of Texas at Austin, Anthropology Department
PROGRAM DESCRIPTION: MA and PhD degrees are offered in Anthropology, with specialization in physical anthropology, including primate anatomy, ecology, evolution, behavior, paleoanthropology, and molecular anthropology.
FACULTY AND THEIR SPECIALTIES: Deborah Bolnick (physical anthropology, molecular anthropology, ancient DNA); Mariah Hopkins (primate conservation, geographic information system (GIS) modeling systems, research in Panama); John Kappelman (physical anthropology, paleobiology, primate evolution, functional morphology, stratigraphy; Africa and Asia); Chris Kirk (physical anthropology, primate evolution, functional anatomy and evolution of sensory systems); Rebecca Lewis (evolution of primate social behavior, power dynamics and power structures, intersexual conflict, social relationships, biological markets, feeding ecology, lemurs); Denné Reed (human evolution, micromammal paleoecology, taphonomy, GIS, remote sensing); and Liza Shapiro (physical anthropology, primate evolution, functional morphology, locomotion).
FOR FURTHER INFORMATION: Graduate Admissions Advisor, Dept. of Anthropology, Univ. of Texas, 1 University Station, C3200, Austin, TX 78712; or see: <>.


* University of Washington, Department of Psychology
PROGRAM DESCRIPTION: The Animal Behavior Program at the University of Washington is dedicated to providing the best possible graduate training including research techniques, theory, and investigative work with animals both in the laboratory and in natural habitats, preserves, and progressive zoos. The program leads to the PhD in Psychology, with special training in animal behavior (including primate social behavior and conservation). It is administered by the core faculty in animal behavior, who are listed below. One of the great assets of this Animal Behavior Program is the interest and competence of faculty in departments other than Psychology. Cordial and cooperative relationships exist with behavior-oriented colleagues in Biology, Anthropology, wildlife science (College of Fisheries and School of Forest Resources), the Center for Conservation Biology, the Neurobiology Program, the Washington National Primate Research Center, and the Center for Human Development and Disabilities. Excellent rapport and research affiliations also exist with the Woodland Park Zoological Gardens, Point Defiance Zoo, the Seattle Aquarium, Northwest Trek, Friday Harbor Laboratory (biology and marine research laboratories), and other colleagues in the greater Puget Sound area.
FACULTY AND THEIR SPECIALTIES: Michael D. Beecher, (animal communication, avian sociobiology and ecology); David P. Barash (sociobiology, behavioral ecology, animal behavior and evolution); Eliot A. Brenowitz (avian behavior, neuroethology, neuroendocrinology, animal communication); Sean O’Donnell (social behavior, especially of insects; evolution of eusociality, particularly division of labor and task allocation; behavioral genetics; and physiology); and Joseph Sisneros (neuroethology; comparative neural bases, anatomy, physiology, function, and modeling of audition). Also available to facilitate student projects are James Ha (social behavior and cognition, behavioral genetics; applied animal behavior; primates, corvids, cetaceans), Randall Kyes (international macaque field site; conservation biology), and Renee Ha (behavior and ecology of corvids; conservation biology), almost all of whom are graduate faculty.
FOR FURTHER INFORMATION: Michael D. Beecher, PhD, Dept. of Psychology, Box 351525, University of Washington, Seattle, WA 98l95-1525 [e-mail: [email protected]].


* Wisconsin National Primate Research Center (WNPRC), University of Wisconsin–Madison, Graduate School, supported by a base operating grant from the National Institutes of Health–National Center for Research Resources
PROGRAM DESCRIPTION: The research program at the WNPRC has opportunities for graduate studies in several areas, especially reproductive and developmental biology, including placental biology and stem cell research, immunogenetics, virology and AIDS vaccine development, veterinary medicine, aging and metabolic disease, psychology and psychobiology, and neurobiology, including Parkinson’s disease research.
Students may conduct research at the WNPRC by enrolling in an appropriate academic department at the University of Wisconsin–Madison and choosing a faculty advisor with WNPRC affiliation. Current faculty have appointments in various departments in the Medical School, College of Letters and Science, School of Veterinary Medicine, and College of Agriculture and Life Sciences, as well as such interdisciplinary programs as the Endocrinology–Reproductive Physiology Program, Interdepartmental Graduate Program in Nutritional Sciences, the Biology of Aging and Age-Related Diseases Training Program, Stem Cell and Regenerative Medicine Center Education and Training Programs, and the Neuroscience Training Program. For information about these departments and programs, potential students should write to The Graduate School, Bascom Hall, UW–Madison, Madison, WI 53706; or visit <>.
FACULTY AND STAFF: The WNPRC supports the work of 125 UW–Madison scientists (27 with base grant support), 112 scientists at other institutions, and 175 staff members.
FOR FURTHER INFORMATION: Donna Paulnock, Interim Director, WNPRC, 1220 Capitol Ct, Madison, WI 53715-1299. Director’s Office and general information: [608-263-3500; fax: 608-265-2067]; or see: <>.


* University of Calgary, Department of Anthropology
PROGRAM DESCRIPTION: Master’s and Doctoral programs are available in primate studies, principally field-based and with a behavioral ecology approach. Both programs require course work, a formal research proposal defense, field research, preparation and defense of a thesis, and candidacy and second-language exams at the doctoral level. The Department has research relationships with various primate centers in the U.S.A.; the Monkey River, Belize, site at which an annual field school is conducted; the Boabeng Fiema Monkey Sanctuary in Ghana at which an annual field school is conducted; Santa Rosa National Park in Costa Rica; and various field sites in Madagascar.
FACULTY AND THEIR SPECIALTIES: Linda Fedigan (life histories, behavioral ecology, and conservation of neotropical primates; field site in Costa Rica); Steig Johnson (behavioral ecology, biogeography, speciation, and conservation biology of lemurs, especially brown lemurs; field sites in Madagascar); Mary Pavelka (social organization, social relationships, aging, and reproduction; field sites for spider and howler monkeys in Belize); and Pascale Sicotte (behavior, social relationships, behavioral ecology, colobines, apes).
FOR FURTHER INFORMATION: Dept. of Anthropology, Univ. of Calgary, Calgary, Alberta T2N 1N4, Canada [e-mail: [email protected], [email protected], [email protected], or [email protected]] or see: <>.

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Judith E. Schrier, Psychology Department, Box 1853, Brown University
Providence, Rhode Island 02912. [401-863-2511; fax: 401-863-1300]
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Cover photograph of a white-handed gibbon(Hylobates lar), taken at the San Diego Zoo by Mark Abbott in 2007

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