Problems in the population and community
Functional morphology, comparative physiology, and biomechanics
Functional morphology, the evolution of the vertebrate locomotor system.
Vertebrate evolution (especially Cenozoic mammals), craniodental functional morphology, mammalian systematics (especially ungulates).
Behavioral ecology: foraging, life history variables, and sexual selection.
Graduate Program Co-Director
Population genetics and evolutionary theory, applied to human genetic variation
Anatomy Course Director/
Department Vice Chair
Species invasions of non-native plants and animals; Evaluating conservation strategies best suited to preserving biodiversity from the novel threats posed by climate change.
Conservation Medicine; Disease Ecology; Biogeography; Global Environmental Change.
Biomechanics, functional morphology, and the evolution of vertebrate limbs.
Life history evolution with an emphasis on Senescence.
Population and community ecology, oceanography, and marine biology.